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Melinda Bennion, Camille Burrus, Kody Callister,
Christian Edwards, Brianna Johnson, Clayton Martinez,
Justin Neighbor, Jason Pietrzak, Mark Ratchford,
Skye Salganek, Martin Schijf, Sarah Siefken, Shawn
Smith, Janice Stroud-Settle, Kevin Wheeler,
Eric Woodhouse, Joe Zant
Zone 3
Zone 1
Zone 2 Zone 4
Z5
0
20
40
60
80
100
120
5
107
35
28
42
65
34
18
28
18 16 16
4
20
13
9
4 6 8 9
NumberofTortoises
309 Adult Tortoises (180-314 mm)
173 juvenile tortoises (38-179 mm)
0
2
4
6
8
10
12
14
16
NumberObserved
Carapace Length (mm)
Males Females
Photo by John Kellam
Determine success of translocation program:
 Spatial distribution of tortoises & sign
 Pop structure, sex ratio, growth, movement
 Density, abundance, survival, mortality, disease
 Compare to adjacent tortoise populations
Photo by John Kellam
Determine success of translocation program:
 Spatial distribution of tortoises & sign
 Pop structure, sex ratio, growth, movement
 Density, abundance, survival, mortality, disease
 Compare to adjacent tortoise populations
Survey Year Transects Km
Surveyed
Tortoises
Observed
0 1987-91 25 42.6 0
1
2003 45 44.1 17
2005 22 42.2 19
2007 34 61.5 13
2009 24 27.9 13
2
2013 17 46.3 15
2015 16 41.8 11
3 2017 30 85.7 45
0
5
10
15
20
25
1987-91 2003-09 2013-15 2017
Density(tortoises/sqkm)
Survey Year
Density in Management Zone 4
0
5
10
15
20
25
1987-91 2003-09 2013-15 2017
Density(tortoises/sqkm)
Survey Period
Density in Management Zone 4
0
100
200
300
400
500
600
1999 2001 2003 2005 2007 2009 2011 2013 2015 2017
AdultTortoiseAbundance(CL>180mm)
Cumulative Total Annual Estimate
5
10
15
20
25
30
35
40
45
1999 2001 2005 2009 2013 2017
Density(tortoise/sqkm)
Monitoring Year
40 Tortoises Observed
6 juveniles: 30 adults
15 ♂: 15 ♀
URTD signs 3/35 (8.6%)
24 previously marked
20 translocated
Mean=11.13 ± 2.38 yr
Range=0.85 to 19.44 yrs
0
2
4
6
8
10
12
14
Male Female Juvenile
Growth(mm/yr)
Translocated Reserve
(Long term growth, Medica et al. 2012)
0
100
200
300
400
500
600
700
800
Z4 (T) Z4(N) Z 2,3,5
TotalDistanceMoved(m)
 Adult survival 0.90 (0.87-0.92)
 Mortality: 27 known deaths (26% roadkill, 18% disease, 3% shot, 3% overturned)
 Adult tortoises appear to have high site fidelity w/in Zone 4
In 2005, fires burned approximately 25% of
tortoise habitat within the Red Cliffs NCA
 Tortoise burrow encounter rate has increased
Observed 0.02 burrows per km in 1987-91 and
0.89 burrows per km from 2003 to 2017.
 Density has increased from very low (undetected) to
13.4 tortoises per sq km (95% CI: 7.5-23.9)
Density in Reserve is 19.6 tortoises per sq km
 Translocated adult tortoises had a significantly higher
growth rate than Reserve adult tortoises.
 Although movement greater for Z4 tortoises,
translocated tortoises have high site fidelity
 Continue to monitor tortoises in Zone 4 and across Reserve
 Continue to translocate tortoises into Zone 4
 Identify management actions (i.e., education, LE presence, signage)
to reduce preventable mortality
 Identify and evaluate future potential translocation sites w/ low density populations

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Status of Translocated Tortoises in Southwest Utah

  • 1.
  • 2. Melinda Bennion, Camille Burrus, Kody Callister, Christian Edwards, Brianna Johnson, Clayton Martinez, Justin Neighbor, Jason Pietrzak, Mark Ratchford, Skye Salganek, Martin Schijf, Sarah Siefken, Shawn Smith, Janice Stroud-Settle, Kevin Wheeler, Eric Woodhouse, Joe Zant
  • 3. Zone 3 Zone 1 Zone 2 Zone 4 Z5
  • 4.
  • 5.
  • 6.
  • 7.
  • 8.
  • 9.
  • 10.
  • 11.
  • 13. 309 Adult Tortoises (180-314 mm) 173 juvenile tortoises (38-179 mm) 0 2 4 6 8 10 12 14 16 NumberObserved Carapace Length (mm) Males Females
  • 14.
  • 15. Photo by John Kellam Determine success of translocation program:  Spatial distribution of tortoises & sign  Pop structure, sex ratio, growth, movement  Density, abundance, survival, mortality, disease  Compare to adjacent tortoise populations
  • 16. Photo by John Kellam Determine success of translocation program:  Spatial distribution of tortoises & sign  Pop structure, sex ratio, growth, movement  Density, abundance, survival, mortality, disease  Compare to adjacent tortoise populations
  • 17.
  • 18.
  • 19. Survey Year Transects Km Surveyed Tortoises Observed 0 1987-91 25 42.6 0 1 2003 45 44.1 17 2005 22 42.2 19 2007 34 61.5 13 2009 24 27.9 13 2 2013 17 46.3 15 2015 16 41.8 11 3 2017 30 85.7 45
  • 20.
  • 21.
  • 22. 0 5 10 15 20 25 1987-91 2003-09 2013-15 2017 Density(tortoises/sqkm) Survey Year Density in Management Zone 4
  • 23. 0 5 10 15 20 25 1987-91 2003-09 2013-15 2017 Density(tortoises/sqkm) Survey Period Density in Management Zone 4
  • 24. 0 100 200 300 400 500 600 1999 2001 2003 2005 2007 2009 2011 2013 2015 2017 AdultTortoiseAbundance(CL>180mm) Cumulative Total Annual Estimate
  • 25. 5 10 15 20 25 30 35 40 45 1999 2001 2005 2009 2013 2017 Density(tortoise/sqkm) Monitoring Year
  • 26.
  • 27. 40 Tortoises Observed 6 juveniles: 30 adults 15 ♂: 15 ♀ URTD signs 3/35 (8.6%) 24 previously marked 20 translocated Mean=11.13 ± 2.38 yr Range=0.85 to 19.44 yrs
  • 28.
  • 29. 0 2 4 6 8 10 12 14 Male Female Juvenile Growth(mm/yr) Translocated Reserve (Long term growth, Medica et al. 2012)
  • 30. 0 100 200 300 400 500 600 700 800 Z4 (T) Z4(N) Z 2,3,5 TotalDistanceMoved(m)  Adult survival 0.90 (0.87-0.92)  Mortality: 27 known deaths (26% roadkill, 18% disease, 3% shot, 3% overturned)  Adult tortoises appear to have high site fidelity w/in Zone 4
  • 31.
  • 32. In 2005, fires burned approximately 25% of tortoise habitat within the Red Cliffs NCA
  • 33.
  • 34.
  • 35.
  • 36.
  • 37.  Tortoise burrow encounter rate has increased Observed 0.02 burrows per km in 1987-91 and 0.89 burrows per km from 2003 to 2017.  Density has increased from very low (undetected) to 13.4 tortoises per sq km (95% CI: 7.5-23.9) Density in Reserve is 19.6 tortoises per sq km  Translocated adult tortoises had a significantly higher growth rate than Reserve adult tortoises.  Although movement greater for Z4 tortoises, translocated tortoises have high site fidelity
  • 38.  Continue to monitor tortoises in Zone 4 and across Reserve  Continue to translocate tortoises into Zone 4  Identify management actions (i.e., education, LE presence, signage) to reduce preventable mortality  Identify and evaluate future potential translocation sites w/ low density populations